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Puzzle - Hint

My first is in horse but not in pony.
My second is in coyote but not in jackal.
My third is in shrew and also in vole.
My fourth is in panda but not in bear.
My last is in baboon but not in monkey.
What am I?

 [Ref: ZCPD] © Kevin Stone

Hint: The first letter can only be H, R, S or E.

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